Doctoral Programs
Biochemistry, Cell Biology, Molecular Biology (BCMB)
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    Doctoral Program
    BCMB Allied Program
    Mouse pancreatic beta cells cultured in a monolayer. Insulin labeled in green. Credit: Rachel Stamateris
  • BCMB 3
    Doctoral Program
    BCMB Allied Program
    Collecting lymphatic vessel shown in green with cell junctions shown with VE-Cadherin staining (red). Credit: Chou Chou
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    Doctoral Program
    BCMB Allied Program
    Whole-scan Maximum Projection of live 4 Days Post Fertilization double Transgenic Casper zebrafish larvae Tg(kdrl:RFP, mpeg1:eGFP), highlighting Endothelium and Macrophages. Credit: Gelashvili, Zaza
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    Doctoral Program
    BCMB Allied Program
    3D reconstructed super resolution image showing mitochondria (pink) surrounding a ruptured (loss of yellow integrity marker) micronucleus (blue, DNA). Credit: Melody Di Bona, PhD
  • BCMB Banner 5
    Doctoral Program
    BCMB Allied Program
    3D reconstruction of super resolution image showing a micronucleus (blue, DNA) surrounded by lamina (green), in which a protein is aggregating (red). Credit: Melody Di Bona, PhD
  • BCMB Banner 8
    Doctoral Program
    BCMB Allied Program
    Maximum Projection of Live 3 Days Post Fertilization Transgenic Casper Zebrafish Larvae, Tg(kdrl: cPLA2-mKate2-p2A-eGFP*-KDEL) expressing marker for Endoplasmic Reticulum and cPLA2 in Endothelial Cells. Credit: Gelashvili, Zaza and Shen, Zhouyang
  • BCMB Banner 9
    Doctoral Program
    BCMB Allied Program
    In vitro-derived 3D mouse gastruloid stained with DAPI (blue), CDX2 (Red), E-CADHERIN (White) and SOX2 (Green). Credit: Mohamad Gatie
  • BCMB Banner 10
    Doctoral Program
    BCMB Allied Program
    Midgestation mouse embryo at E8.75 stained with Uncx (Yellow), Nkx2.1 (Green), Ripply3 (Red), Cldn6 (White) and DAPI (Blue) using mRNA-multiplex HCR. Credit: Mohamad Gatie
  • BCMB Banner 13
    Doctoral Program
    BCMB Allied Program
    Alpha-synuclein binding site (dot plot) mapped onto the surface of Rab3a. Credit: Guohua Lv
  • BCMB Banner 13
    Doctoral Program
    BCMB Allied Program
    Human NEXT helicase complex bound to RNA. Credit: Chris Lima
  • BCMB Banner 14
    Doctoral Program
    BCMB Allied Program
    Human RNA exosome unwinding RNA. Credit: Chris Lima

    Our Students

    Entering Classes

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    2025
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    2024
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    2023
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    2022
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    2021
    2020
    2020
    Current Cohort

    Student

    Year Entered

    Program

    Year in Program

    Major Sponsor

    Regan, Samantha Brooke

    AY 2018-2019

    Cell & Developmental Biology

    G8

    Maria Jasin

    Cheng, Shuyuan

    AY 2019-2020

    Cell & Developmental Biology

    G7

    Alex Kentsis

    Durani, Vidushi

    AY 2019-2020

    Cell & Developmental Biology

    G7

    Charles Rudin

    Gallegos, Valerie

    AY 2018-2019

    Molecular Biology

    G7

    Olivier Elemento

    Liu, Jiacheng

    AY 2019-2020

    Biochemistry & Structural Biology

    G7

    Christopher Lima

    Perder, Bjoern

    AY 2019-2020

    Cell & Developmental Biology

    G7

    Jingli Cao

    Almodovar, Nickolas

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Benjamin Kleaveland

    Bednor, Lauren

    AY 2020-2021

    Molecular Biology

    G6

    Beate Schwer

    Chu, Kevan

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Lukas Dow

    Han, Qixiu

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Joao Xavier

    Jiang, Chenyang

    AY 2020-2021

    Molecular Biology

    G6

    Steven Josefowicz

    Kapoor, Aakanksha Rajiv

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Vivek Mittal

    Lau, Aristotle Cheng

    AY 2020-2021

    Biochemistry & Structural Biology

    G6

    John Blenis

    Lee, Uk Jin

    AY 2020-2021

    Molecular Biology

    G6

    Effie Apostolou

    Meng, Yinuo

    AY 2020-2021

    Molecular Biology

    G6

    Rohit Chandwani

    Owiredu, Jude Nana Yaw

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Christopher Barbieri

    Pant, Swati

    AY 2020-2021

    Biochemistry & Structural Biology

    G6

    Stephen Long

    Rajkumar, Sandy Madonna

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Yicheng Long

    Shim, Abraham C

    AY 2020-2021

    Molecular Biology

    G6

    John Maciejowski

    Sisso, Whitney Juliet

    AY 2020-2021

    Cell & Developmental Biology

    G6

    Lukas Dow

    Striepen, Josefine Ursel

    AY 2020-2021

    Molecular Biology

    G6

    John Maciejowski

    Upadhya, Samsara Wynona

    AY 2020-2021

    Biochemistry & Structural Biology

    G6

    Tobias Meyer

    Chwat-Edelstein, Tzippora

    AY 2021-2022

    Molecular Biology

    G5

    Xiaolan Zhao

    DeBitetto, Emily

    AY 2021-2022

    Molecular Biology

    G5

    Agnel Sfeir

    Edgar, Claudia L

    AY 2021-2022

    Molecular Biology

    G5

    Jessica Tyler

    Fall, William Bellizzi

    AY 2021-2022

    Molecular Biology

    G5

    Rohit Chandwani

    Guerra, Yanira

    AY 2021-2022

    Cell & Developmental Biology

    G5

    Melissa Davis

    Hadjimichael, Evi

    AY 2021-2022

    Molecular Biology

    G5

    Kirk Deitsch

    Kasliwal, Kritika

    AY 2021-2022

    Cell & Developmental Biology

    G5

    Matthias Stadtfeld

    Klein, Alison D

    AY 2021-2022

    Cell & Developmental Biology

    G5

    Michael Overholtzer

    Liberatore, Katarina

    AY 2021-2022

    Cell & Developmental Biology

    G5

    Santosha Vardhana

    Miller-Browne, Victoria Nicole

    AY 2021-2022

    Molecular Biology

    G5

    Xiaolan Zhao

    Mishra, Tanmay

    AY 2021-2022

    Cell & Developmental Biology

    G5

    Andrea Ventura

    Ohnstad, Amelia

    AY 2021-2022

    Cell & Developmental Biology

    G5

    Ashley Laughney

    Shah, Hina

    AY 2021-2022

    Molecular Biology

    G5

    Agnel Sfeir

    Turner, Briana Simone

    AY 2021-2022

    Biochemistry & Structural Biology

    G5

    Kayvan Keshari

    Balic, Zerina

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Thomas Vierbuchen

    Chiu, Yu-Hui

    AY 2022-2023

    Biochemistry & Structural Biology

    G4

    Christopher Lima

    Cui, Ruobing

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Craig Thompson

    Duren-Lubanski, Alyssa

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Olivier Elemento

    Goetzler, Emily Margalit

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Matthias Stadtfeld

    Goldberg, Hailey Victoria

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Scott Lowe

    Johnsson, Emma

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Tobias Walther

    Kops, Max

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Lukas Dow

    Lee, Young Sun

    AY 2021-2022

    MD-PhD Cell & Developmental Biology

    G4

    Charles Sawyers

    Lin, Qianqian

    AY 2022-2023

    Molecular Biology

    G4

    David Scheinberg

    Londoner, Rebecca

    AY 2022-2023

    Biochemistry & Structural Biology

    G4

    Alexander Gitlin

    Manrique, Cassandra Shea

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Mary Baylies

    Montero, Angela Margaret

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Lydia Finley

    Nathoo, Isaac Abdalla

    AY 2021-2022

    MD-PhD Biochemistry & Structural Biology

    G4

    Marcus Goncalves

    Peterson, Gabrielle M

    AY 2022-2023

    Biochemistry & Structural Biology

    G4

    Jeremy Dittman

    Ramsamooj, Shakti

    AY 2022-2023

    Biochemistry & Structural Biology

    G4

    David Simon

    Selvaraj, Monica

    AY 2022-2023

    Molecular Biology

    G4

    Agnel Sfeir

    Yin, Hang

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Olivier Elemento

    Zheng, Jian

    AY 2022-2023

    Molecular Biology

    G4

    Xiaolan Zhao

    Zhong, Yaofeng

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Christine Mayr

    Zhu Jiang, Xin Yu

    AY 2022-2023

    Cell & Developmental Biology

    G4

    Eric Lai

    Zou, Mengyang

    AY 2022-2023

    Biochemistry & Structural Biology

    G4

    Tobias Meyer

    Barnard, Emilee Frances

    AY 2023-2024

    Molecular Biology

    G3

    Michael Glickman

    Calia, Giuliana Pia

    AY 2023-2024

    Molecular Biology

    G3

    Effie Apostolou

    Carpen, Peyton Lee

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Matthew Greenblatt

    Cohen-Abeles, Luke

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Tobias Walther

    Crater, Jacqueline M

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Andrew Intlekofer

    Earsley, Alexander (Alex)

    AY 2023-2024

    Biochemistry & Structural Biology

    G3

    Olga Boudker

    Hombalkar, Saloni Kedar

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Michael Overholtzer

    Jia, Su (Alex)

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Maria Jasin

    Jiang, Yi-Zhen

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Maria Jasin

    Kelemen, Yosip

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Shahin Rafii

    Ogrean, Riley Olivia

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Jessica Sheu-Gruttadauria

    Pajuelo, Lourdes

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Daniel Bachovchin

    Pramod, Sneha

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Mara Sherman

    Ravichandran Damodaran, Jeyaram (Jey)

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Tuomas Tammela

    Sandborg (Sandborg), Holly Kay

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Mara Sherman

    Sidhu, Asha V

    AY 2023-2024

    Biochemistry & Structural Biology

    G3

    Tobias Meyer

    Su, Rebecca

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Ashley Laughney

    Ton, Carolyn

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Jennifer Zallen

    Vijayakumar (Vijayakumar), Ragavi

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Thomas Vierbuchen

    Wu, Rebecca Jasmine

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Todd Evans

    Yavid, Jennifer (Jen)

    AY 2023-2024

    Molecular Biology

    G3

    Iestyn Whitehouse

    Zhao (Zhao), Jiahui

    AY 2023-2024

    Cell & Developmental Biology

    G3

    Danwei Huangfu

    Baasan, Dulguun

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Jayanta Chaudhuri

    Bahadur, Sher

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Lukas Dow

    Bailey (Bailey), Benton Montgomery

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Agnel Sfeir

    Bohn, Eliana Harmony

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Effie Apostolou

    Collette, Abbie Marie

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Melinda Diver

    DeFrancisco, Sarah

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Mary Baylies

    Huang (Huang), Yuting (Elisa)

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Xuejun Jiang

    Kim, Jung Eun

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Maria Jasin

    Kim, Minseo

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Scott Lowe

    Li, Joshua Deyuan (Josh)

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Asmin Tulpule

    Liu, Chuyan (Katerina)

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Olivier Elemento

    Lu, Allen Wanglong

    AY 2023-2024

    MD-PhD Biochemistry & Structural Biology

    G2

    Melinda Diver

    Marchese, William (Jack)

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Matthias Stadtfeld

    Monge, Molly Maria-Mckenna

    AY 2023-2024

    MD-PhD Molecular Biology

    G2

    Naama Aviram

    Pacini, Cole

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Alexander Gitlin

    Raichur, Ruth

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Gabrielle Rizzuto

    Smith, Eleanor Josephine

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Dirk Remus

    Sun (Sun), Gary Lin

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Yicheng Long

    Volckova, Eva

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Jessica Sheu-Gruttadauria

    Wu, Elaine

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Junhong Choi

    Xia, Wenxin (Vincent)

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Scott Keeney

    Zhang, Jenny Zhen Yao

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Ashley Laughney

    Zhang, Qingyue

    AY 2024-2025

    Biochemistry, Cell & Molecular Biology

    G2

    Tobias Walther

    Caile, Nolan Thomas

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Cirulli, Alessandro

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Fan, Xiaoyu

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Jia (Jia), Hanyi

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Lin, Sabrina

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Liu, Hannah Ming-hui

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Neaher, Sofia Marina

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Schor, Gail Hedva (Gili)

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Scott, William Shem (Shem)

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Xiao, Leo

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Zhang, Yongyan

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Zhao, Zihan

    AY 2025-2026

    Biochemistry, Cell & Molecular Biology

    G1

    Student Outcomes

    Since 2017 we have graduated 167 doctoral students. Below we list just a few of these alumni to illustrate the kinds of paths that our doctoral students have taken. 

    Alumni

    Current Position

    Year of PhD

    Emily Costa, PhD

    Civic Science Associate, Rita Allen Foundation

    2024

    Bryce Delgado, PhD

    Postdoctoral Fellow, Genentech

    2023

    Helene Jahn, PhD

    Postdoctoral Fellow, Oregon Health & Science University

    2023

    Christopher Nötzel, PhD

    Postdoctoral Fellow, University of Pennsylvania

    2022

    Xiang Li, PhD

    Postdoctoral Fellow, Harvard Medical School/Boston Children’s Hospital

    2022

    Joshua Weiss, MD, PhD

    Resident in Internal Medicine, Weill Cornell Medicine

    2021

    Tapojyoti Das, PhD

    Postdoctoral Fellow, St. Jude Children’s Research Hospital

    2021

    Pradeep Ramalingam, PhD

    Assistant Professor, University of Florida

    2020

    Nevin Yusufova, PhD

    Assoc. Director, Translational Research, AstraZeneca

    2020

    Mridula Balakrishnan, PhD

    Postdoctoral Fellow, Max Planck Institute for Heart and Lung Research

    2019

    Maria Bustillo, PhD

    Postdoctoral Fellow, NYU School of Medicine

    2019

    Diana Wang, PhD

    Dermatology Resident, UCLA

    2018

    Benjamin Whitlock, PhD

    Scientific Lead, Loxo Oncology at Lilly

    2018

    Faranak Fattahi, PhD

    Assistant Professor, UCSF

    2017

    Christopher Fiorese, PhD

    Postdoctoral fellow, Cal Tech

    2017

    Contact Information Program Coordinator:
    Vanessa Dodson
    Program Coordinator:
    Denise Jenkins
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