Doctoral Programs
Immunology & Microbial Pathogenesis
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    Doctoral Progam
    Immunology & Microbial Pathogenesis
    Macrophages infected with Mycobacterium tuberculosis (green) which have been stained for inducible nitric oxide synthase (iNOS) shown in red, an enzyme that catalyzes the production of nitric oxide from L-arginine to restrict bacterial growth. The nucleus of the infected cells is shown in blue. Credit: Miljan Stupar
  • IMP Sonnenberg
    Doctoral Program
    Immunology & Microbial Pathogenesis
    Intestinal inflammation with epithelial cells (green) exhibiting cell death (marked by cleaved caspase 3 in red). Credit: Dr. G. Sonnenberg.
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    Doctoral Program
    Immunology & Microbial Pathogenesis
    Innate lymphocytes (red and green) interacting with microbiota-specific regulatory T cells (blue, green and cyan) in lymphoid tissues. Credit: Dr. G. Sonnenberg.
  • IMP Banner 5
    Doctoral Program
    Immunology & Microbial Pathogenesis
    Mycobacterium smegmatis elongates predominantly from one of the two cells poles. Credit: of Drs. H. Botella and S. Ehrt.

    Student Cohorts & Career Outcomes

    Overview

    Below is a representation of IMP student cohorts organized by entering year, including their thesis lab, thesis project, undergraduate institution, and awards.

    2025
    Entering Class of 2025

    Name

    Undergraduate Institution

    Distinctions

    Noelia Boldizsar

    Duke University

    MD-PhD

    Ottavia Bourdain

    Hunter College

    Isabelle Bursulaya

    University of California - Los Angeles

    Chimere Ezeiruaku

    University of Massachusetts - Amherst

    Qinyan Feng

    University of California - Los Angeles

    Joe Hochberg

    University of Pennsylvania

    Rachel Lorenc

    Johns Hopkins University

    MD-PhD

    George Payne

    University of Virginia

    Lisette Peres-Tintin

    Boston University

    Michelle Wallerius

    University of California - Los Angeles

    Elina Wells

    University of California - Berkeley

    Jenny Wang

    Cornell University

    MD-PhD

    Temima Yellin

    Yeshiva University

    Nuzat Zaman

    Cornell University

    2024
    Entering Class of 2024

    Name

    Undergraduate Institution

    Kasra Askari

    University of California - Los Angeles

    Isabele Johnson

    Haverford College

    Qi Jun Andrew Lim

    Imperial College of London

    Riley Outen

    Haverford College

    Cristian Ovies

    DePaul University

    Catherine Somoza

    Cornell University

    2023
    Entering Class of 2023

    Name

    Thesis Lab

    Undergraduate Institution

    Wiam (Weema) Abdalla Mo Madani

    Juan Cubillos-Ruiz

    St. Olaf College

    Emre Davud Cardakli

    Randy Longman

    Duke University

    Charles Carr

    Nick Collins

    University of Toronto

    Aiquan Chang

    Sallie Permar

    University of California - Berkeley

    Ana Campos Codo

    Justin Perry

    Sao Paulo State University

    Sofia Dimitriadoy

    Andrea Schietinger

    Princeton University

    Michael Downey

    Alexander Rudensky

    Cornell University

    Muxue (Snow) Du

    Steven Josefowicz

    Muhlenberg College

    Sherry Fan

    Joseph Sun

    University of Maryland - College Park

    Mergim Gjonbalaj

    Gabi Rizzuto

    Stony Brook University

    Gregory Gladkov

    Brad Jones

    University of California - Berkeley

    Aya Halawai

    Katharine Hsu

    Massachusetts Institute of Technology

    Victoria Rose Lawless

    Steven Josefowicz

    University of Chicago

    Esther Lee

    Morgan Huse

    Bowdoin College

    Fitty Liu

    Tobias Hohl

    University of California - Berkeley

    Tomi McGuire

    Joseph Sun

    Northeastern University

    Itzayana Miller

    Brad Jones

    Davidson College

    Daniel Min

    Juan Cubillos-Ruiz

    University of Florida

    Connor Smith

    Brad Jones

    University of Virginia

    Eric Yiyang Wang

    Alexander Rudensky

    Yale University

    Claire Wingert

    Lionel Ivashkiv

    University of California - Berkeley

    Yulin Zhou

    Melody Zeng

    University of Toronto

    David Zurowski

    Rachel Niec

    Drexel University

    2022
    Entering Class of 2022

    Name

    Thesis Lab

    Thesis Project

    Undergraduate Institution

    Allison Catalano

    Georgetown University

    Yasmine Issah

    Sallie Permar

    University of Pennsylvania

    Suparna Kumar

    Theresa Lu

    University of Mumbai

    Day Li

    Craig Thompson

    Peking University

    Yun Lo

    Chrysothemis Brown

    Regulation of intestinal tolerance by Thetis cells

    National Yang-Ming University

    Ian McBain

    Andrea Schietinger

    The role of CD4 T cells in driving autoimmune CD8 T cell differentiation and function in type 1 diabetes

    New York University Abu Dhabi

    Joshua Griffin

    Alexander Gitlin

    Decoding the spatial nature of TNF-mediated cell death in intestinal pathology

    Princeton University

    Jonah Pierce

    Marcel Van den Brink

    University of California, Santa Cruz

    Purnima Ravisankar

    Melody Zeng

    Anna University

    Cristine (Nicole) Salinas

    Tobias Hohl

    Innate immunity during gastrointestinal colonization with Candida albicans

    University of Texas at Arlington

    Shane Vedova

    Richard Brad Jones

    of California, Berkeley

    Daniel Wong

    Melody Zeng

    University of British Columbia

    Sufia Bakshi

    Andrea Schietinger

    2021
    Entering Class of 2021

    Name

    Thesis Lab

    Graduation Year

    Thesis Project

    Undergraduate Institution

    Awards

    Giorgi Beroshvili

    Alexander Rudensky

    A nuanced requirement for Foxp3 for Treg cell function

    Ivane Javakhishvili State University

    Hannah Carrow

    Melody Zeng

    Defining the role of bacterial sphingolipids in intestinal immune regulation

    Northeastern University

    Elizabeth Emanuel

    David Artis

    Macalester College

    Aazam Ghelani

    Alexander Rudensky

    Combinatorial role of Treg effector molecule in Treg cell function

    Savitribai Phule Pune University

    Alberto Herrera

    Brad Jones

    Multimodal characterization of persistence profiles in HIV-expressing cells and authentic HIV-reservoir harboring clones

    Seton Hall University - South Orange

    Jee Hye Kang

    Roberta Zappasodi

    Exploring the Effect of Anti-CTLA-4 Variants on Anti-Tumor Immune Response in Cancers with Different Glycolytic Capacity

    University of California-Berkeley

    Cheryl Mai

    Alexander Rudensky

    Yale University

    Dean Matthews

    Gretchen Diehl

    Regulation of Immune Development by Commensal-Specific IgG

    University of Pittsburgh

    Nazia Sakila

    Kyu Rhee

    2025

    Columbia University

    Sophia Qu

    Gregory Sonnenberg

    Cornell University

    NSF Award Recipient

    Baila Schochet

    Inez Rogatsky

    CUNY - Hunter College

    Brittany Robaina-Caicedo

    Jayanta Chaudhuri

    University of Central Florida

    NSF Award Recipient

    2020
    Entering Class of 2020

    Name

    Thesis Lab

    Graduation Year

    Thesis Project

    Undergraduate Institution

    Awards

    Mamadou Bah

    Jedd Wolchok

    Investigating dynamic IDO-Kyn-AhR Pathway Induced Tumor Immunosuppression

    New Jersey City University

    Michael Bale

    Steven Josefowicz

    Chromatin dynamics throughout immune cell development and stimulation

    University of Kentucky

    Ryann Callaghan

    Gretchen Diehl

    Defining how intestinal microbes promote colorectal cancer through modulation of macrophage function

    Drew University

    Amanda Chen

    Gretchen Diehl

    Investigating the role of the microbiome in mononuclear phagocyte differentiation and function

    University of California-Davis

    Logan Fisher

    Chrysothemis Brown

    Uncovering Heterogeneity of CCR7+ Dendritic Cells

    University of California-Santa Cruz

    Joseph Gladstone

    Gregory Sonnenberg

    Examining innate like lymphocytes in intestinal health, inflammation, and homeostasis

    Northeastern University

    Dasom Kim

    Gretchen Diehl

    2025

    Microbiota-induced host regulatory networks to limit microbiota-specific T cell responses

    Brigham Young University

    Noemi Linden

    R. Brad Jones

    Characterization of T Cell-inherent and Virus-conferred Factors Interfering with Immune Clearance of HIV

    Jacobs University

    Jez Marston

    Ethel Cesarman

    2024

    Identifying Endogenous Retroviral Factors in Viral Lymphomagenesis

    Yale University

    Svetlana Miakicheva

    Andrea Schietinger

    Decoding and Reprogramming Autoimmune Stem-like T cells in Type 1 Diabetes

    University of California-Davis

    Tanmana Mitra

    Santosha Vardhana

    Regulation of T cell metabolism by antigen avidity

    Indian Institute of Technology

    Mark Owyong

    Joseph Sun

    Elucidating the role of SATB1 in NK cell development and antiviral host immunity

    University of California-Berkeley

    F31 Award Recipient

    Yoselin Paucar

    Chrysothemis Brown

    John Jay College of Criminal Justice

    HHMI Gilliam Award Recipient

    Gayathri Shibu

    Chrysothemis Brown

    Role of peripheral AIRE in self tolerance

    Vit University

    Student Career Outcomes

    IMP graduates have successfully embarked upon a variety of careers, including as academic faculty positions and researchers in industry. Below is a representation of some of our IMP alumni, their current positions, and the lab they trained in as a graduate student at WCGS from over the years: 

    Name

    Current Position

    Lab

    John MacMicking, PhD

    Professor of Microbial Pathogenesis and Immunobiology, HHMI Investigator

    Carl Nathan

    Xiaoyu Hu, PhD

    Professor, Tsinghua University (China)

    Lionel Ivashkiv

    Cynthia Leifer, PhD

    Professor, Cornell University

    Carl Nathan

    Utthara Nayar , PhD

    Professor, Johns Hopkins University

    Ethel Cesarman

    Alexander Ploss, PhD

    Harry C. Wiess Professor in the Life Sciences, Princeton University

    Eric Pamer (Now at Univ. Of Chicago)

    Michael Shiloh, PhD

    Associate Professor, UT Southwestern

    Carl Nathan

    Aimee Beaulieu, PhD

    Assistant Professor & Chancelor Scholar, Rutgers Medical School

    Carl Nathan 

    Minkyung Song, PhD

    Assistant Professor, Sungkyunkwan Univ (Korea)

    Juan Cubillos-Ruiz

    Katherine Hisert, M.D, PhD

    Assistant Professor at National Jewish Health

    Carl Nathan

    Robert Samstein, M.D., PhD

    Assistant Professor, Radiation Oncologist, Icahn School of Medicine at Mount Sinai

    Alexander Rudensky

    Ruth Franklin, PhD

    Assistant Prof., Stem Cell and Regenerative Biology, Harvard University

    Ming Li

    Sungho Park, PhD

    Assistant Professor, UNIST (Korea)

    Lionel Ivashkiv

    Rachel Gottschalk, PhD

    Assistant Prof., Integrative Systems Biology, Dept. of Immunology, Univ. of Pittsburgh

    Jim Allison (now at M.D. Anderson Cancer Center)

    Kang Chen, PhD

    Professor, Wayne State University

    Andrea Cerutti (now at Mount Sinai)

    Joris van der Veeken, PhD

    Assistant Professor, IMP (Austria)

    Alexander Rudensky

    Clarissa Campbell, PhD

    Assistant Professor, Center for Molecular Med (Austria)

    Alexander Rudensky

    Jen Oyler, PhD

    Assistant Professor of Systems Biology, Harvard Medical School

    Gregoire Altan-Bonnet(now at NIH)

    Kyuho Kang, PhD

    Assistant Professor, Chungbuk University (Korea)

    Lionel Ivashkiv

    Virginia Pedicord, PhD

    Assistant Professor, University of Cambridge (UK)

    Jim Allison (now at M.D. Anderson Cancer Center)

    Jennifer Small Saunders, M.D., PhD

    Assistant Professor of Medicine, Columbia University

    Sabine Ehrt

    Sheetal Gandotra, PhD

    Assistant Professor, CSIR-IGIB New Delhi (India)

    Sabine Ehrt

    Gregory St John, PhD

    Therapeutic Area Lead, Dermatology US Medical Affairs at AbbVie

    Carl Nathan

    Omar Vandal, PhD

    Principal Technologist, Accelerator, Gates Foundation

    Sabine Ehrt

    Jenny Tzeng

    Associate Director, Bioscience Immunology, AstraZeneca

    Theresa Lu

    Younghwa Kim, M.D., PhD

    Sr Director Clinical Science, Early Clinical Oncology, AstraZeneca

    Carl Nathan

    Jessica Schneider, PhD

    Chief Scientific Officer, Corundum Systems Biology

    Michael Glickman

    Shuangping Shi, PhD

    Associate Vice President, Head of Biologics Process Research & Development, Merck

    Sabine Ehrt

    Kan Lin, PhD

    Kan Lin, PhD Managing Director, Hankang Capital (China)

    Sabine Ehrt 

    Joon Seok Park, PhD

    Assistant Professor, University of Chicago

    Alexander Tarakhovsky & Alexander Rudensky

    Rachel Niec, M.D. PhD

    Assistant Attending Physician, Assistant Professor, Memorial Sloan Kettering Cancer Center & Weill Cornell Graduate School

    Alexander Rudensky

    Gabriel Loeb, M.D., PhD

    Assistant Professor of Medicine, UCSF

    Alexander Rudensky

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